combining form of prec., as spinuloso-ciliate, -dentate, -denticulate, -granulate, -serrate.

1

1833.  Hooker, in Smith’s Eng. Flora, V. I. 121. The margins … more or less spinuloso-dentate. Ibid., 124. The upper lobes … spinuloso serrate.

2

1846.  Dana, Zooph. (1848), 162. Corallum having a subelliptical aperture, sides … spinuloso-denticulate. Ibid. (1852), Crust., I. 186. Fourth, fifth and especially sixth joint spinuloso-granulate above.

3

1866.  Treas. Bot., 1084/2. Spinuloso-ciliate, ciliated with fine spines.

4