combining form of prec., as spinuloso-ciliate, -dentate, -denticulate, -granulate, -serrate.
1833. Hooker, in Smiths Eng. Flora, V. I. 121. The margins more or less spinuloso-dentate. Ibid., 124. The upper lobes spinuloso serrate.
1846. Dana, Zooph. (1848), 162. Corallum having a subelliptical aperture, sides spinuloso-denticulate. Ibid. (1852), Crust., I. 186. Fourth, fifth and especially sixth joint spinuloso-granulate above.
1866. Treas. Bot., 1084/2. Spinuloso-ciliate, ciliated with fine spines.